He was born in 1939 in Ponta Delgada and is Professor Emeritus. He holds a Degree in Medicine (1963) from the Faculty of Medicine of the University of Coimbra and in that same year he began working there as a Second Assistant in the subject of Histology and Embryology. He holds a Doctorate in Histology and Embryology (1987) and has been an Associate Professor since 2001.
In 1976 he became the Director of the Professor Doutor Costa Simões Histology and Embryology Institute at the Faculty of Medicine of the University of Coimbra. He directs the histology and embryology classes at the Faculty of Medicine of the University of Coimbra. He was a member of the Advisory Board in the areas of the Health Sciences at the National Institute for Scientific Research. He was a founding member of the Cellular Biology Centre at the University of Coimbra. He founded the Centre for Histophysiology, Experimental Pathology and Developmental Biology (FCT). He was the Director of the Electronic Microscopy Lab at the Faculty of Medicine of the University of Coimbra. He was the President of the Portuguese Society of Electronic Microscopy.
Books published in Coimbra University Press: